Research Group of Plant and Vegetation Ecology, Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Antwerp, Belgium

Laboratory of Plant Ecology, Department of Applied Ecology and Environmental Biology, Ghent University, Coupure Links 653, 9000 Ghent, Belgium

Institute of Forest Ecology, Mendel University of Agriculture and Forestry, Zemědělská 3, 61300 Brno, Czech Republic

Research Group of Plant and Vegetation Ecology, Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Antwerp, Belgium

Research Group of Plant and Vegetation Ecology, Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Antwerp, Belgium

Research Institute for Nature and Forest, Ministry of the Flemish Community, Gaverstraat 4, 9500 Geraardsbergen, Belgium

Laboratory of Plant Ecology, Department of Applied Ecology and Environmental Biology, Ghent University, Coupure Links 653, 9000 Ghent, Belgium

Institute of Forest Ecology, Mendel University of Agriculture and Forestry, Zemědělská 3, 61300 Brno, Czech Republic

Research Group of Plant and Vegetation Ecology, Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Antwerp, Belgium

Research Group of Plant and Vegetation Ecology, Department of Biology, University of Antwerp, Universiteitsplein 1, 2610 Antwerp, Belgium